Education

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Teaching

Fall Semester

Fundamentals of Osteopathic Medicine

Neuromusculoskeletal Medicine

Growth and Development (Elective Course)

Spring Semester

Cardiovascular, Respiratory and Renal Medicine

Gastrointestinal Medicine

Reproductive Medicine

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Research Interests

Primate Evolution

I am interested in how the Primate Order arose and evolved, principally as seen through the fossil record. My direct field research experience in this area has been with South American primates, mostly in Brazil. My next field project involves paleontological survey in south Asia.

Primate Life History

I am interested in the tradeoffs between neural growth and somatic growth in primates, particularly in terms of how primates have occupied niche spaces in competition with more specialized vertebrates. I believe that the ecological distribution of primates around the world can be explained largely on the basis of a species’ relative commitment to a “cognitive” niche.

History of Science

I am interested in how questions about evolution, particularly primate and human evolution, are framed, executed and interpreted scientifically. I try to account for current trends or consensus opinions in terms of how data have accumulated over time. I advocate for the value of inductive reasoning in scientific inquiry.

Medical Education

I am interested in the pedagogy of basic science medical education. Toward this end my scholarly efforts have focused on the publication of an anatomy and embryology textbook in February of 2007 and a guide to medical school published in 2009.

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Publications

Books

2009 Hartwig, WC Med School Rx: Getting In, Getting Through and Getting On With Doctoring. New York: Kaplan Publishing.

2007 Hartwig, WC Fundamental Anatomy. Lippincott, Williams & Wilkins

2002 Hartwig, WC (editor) The Primate Fossil Record. Cambridge: Cambridge University Press.


Recent Articles and Chapters

In press Hartwig, WC Introduction. In Ecology and Conservation of Sakis, Titis and Uacaris. (Barnett and Veiga, eds.) Cambridge: Cambridge University Press.

In press Hartwig, WC Primate Evolution and Taxonomy. In Primates in Perspective, 2nd Edition. (Campbell, Panger, Fuentes, MacKinnon and Bearder, eds.) Oxford: Oxford University Press.

2008 Hartwig, WC Evolution on the road less traveled. Review of Primate Evolution, by Susan Cachel. American Journal of Primatology 70:1003-1005.

2008 Rosenberger, AL, Halenar, L, Cooke, S, and Hartwig, WC Morphology and evolution of the spider monkey, genus Ateles. In Spider Monkeys: Behavior, Encology and Evolution of the Genus Ateles (Campbell CJ, ed.). Cambridge: Cambridge University Press, pp. 19-49.

2007 Hartwig, WC Obituary: F. Clark Howell, Evolutionary Anthropology 16:81-82.

2007 Hartwig, WC Selection and the outer self. Review of Skin: A Natural History, by Nina G. Jablonski. Journal of Mammalian Evolution 14:200-201.

2006 Hartwig, WC Primate Evolution. In Primates in Perspective (Campbell, Panger, Fuentes, MacKinnon and Bearder, eds.). Oxford University Press, pp. 11-22.

2005 Hartwig, WC Implications of molecular and morphological data for understanding ateline phylogeny. International Journal of Primatology 26:999-1015.

2005 Hartwig, WC and Rosenberger, AL New World monkeys. In Encyclopedia of Life Sciences. London: Macmillan. (www.els.net/els/els/els/index.html?

2005 Hartwig, WC and Rosenberger, AL Primates. In Encyclopedia of Life Sciences. London: Macmillan. /els/els/els/index.html?sessionid=e8808eb7ecf71d4f)

2004 Hartwig, WC Review of Life’s Solution by Simon Conway Morris. American Journal of Human Biology 16:496-497


Prior Articles

2002 Whitaker, JM, Williams, T, Rousseau, L., Rowan, RA, and Hartwig, WC A scoring system for estimating age in the foot skeleton. American Journal of Physical Anthropology 118:385-392.

2002 Hartwig, WC Introduction to the Primate Fossil Record. In The Primate Fossil Record (WC Hartwig, ed.), pp. 1-4. Cambridge: Cambridge University Press.

2002 Hartwig, WC and Meldrum, DJ Miocene platyrrhines of the northern Neotropics. In The Primate Fossil Record (WC Hartwig, ed.), pp. 175-188. Cambridge: Cambridge University Press.

2001 Hartwig, WC and Rosenberger, AL Old World monkeys. In Encyclopedia of Life Sciences. London: Macmillan. www.els.net/els/els/els/index.html? sessionid=e8808eb7ecf71d4f)

2001 Hartwig, WC Fossils, footprints and foragers: bipedalism evolving. Evolutionary Anthropology 10:3-4.

1999 Chamberlain JG and Hartwig WC Thomas Kuhn and paleoanthropology. Evolutionary Anthropology 8:42-44.

1999 Hartwig, WC Primate evolution. In The Nonhuman Primates (P Dolhinow & A Fuentes, eds). New York: Mayfield Press, pp.10-17.

1998 Masterson, TJ and Hartwig, WC Degrees of sexual dimorphism in Cebus and other New World monkeys. American Journal of Physical Anthropology 107:243-256.

1998 Hartwig, WC and Doneski, KF Evolution of the hominid hand and tool making behavior. American Journal of Physical Anthropology 106:401-404.

1998 Hartwig, WC Language origins and diversification. Evolutionary Anthropology 6:39-40.

1997 Fleagle, JG and Hartwig, WC Paleoprimatology. In History of Physical Anthropology: An Encyclopedia (F. Spencer, ed.). New York: Garland, pp.796-810.

1997 Cartelle, C and Hartwig, WC Primatologia de Sudamerica: los grandes platirrinos. Investigacion y Ciencia (April):30-32.

1997 Hartwig, WC Squirrel monkeys as animal models of pelvic relaxation. American Journal of Obstetrics and Gynecology 177:489-490.

1996 Hartwig, WC Perinatal life history traits in New World monkeys. American Journal of Primatology 40:99-130.

1996 Hartwig, WC and Cartelle, C A complete skeleton of the giant South American primate Protopithecus. Nature 381:307-311.

1996 Cartelle, C and Hartwig, WC A new extinct primate among the Pleistocene megafauna of Bahia, Brazil. Proceedings of the National Academy of Sciences 93:6405-6409.

1996 Hartwig, WC; Rosenberger, AL; Garber, PW; Norconk, MA On ateline evolution. In MA Norconck, AL Rosenberger and P Garber (eds.): Adaptive Radiations of Neotropical Primates. New York: Plenum Press, pp.427-431.

1996 Hartwig, WC Physical anthropology today and tomorrow. Evolutionary Anthropology 4:160

1996 Cartelle, C and Hartwig, WC Updating the two Pleistocene primates from Bahia, Brasil. Neotropical Primates 4:46-48.

1996 Hartwig, WC A model human biology program. Evolutionary Anthropology 5:1-2.

1996 Cartelle, C and Hartwig, WC Macacos sul-americanos: descoberta importante muda a história. Ciência Hoje 21:30-37.

1995 Hartwig, WC A giant New World monkey from the Pleistocene of Brazil. Journal of Human Evolution 28:189-196.

1995 Hartwig, WC Effect of life history on the squirrel monkey (Platyrrhini, Saimiri) cranium. American Journal of Physical Anthropology 97:435-449.

1995 Hartwig, WC Protopithecus: Rediscovering the first fossil primate. History and Philosophy of the Life Sciences 17:447-460.

1994 Hartwig, WC Patterns, puzzles and perspectives on platyrrhine origins. In Integrative Paths to the Past: Paleoanthropological Advances in Honor of F. Clark Howell (RS Corruccini and RL Ciochon, eds.). New York: Prentice-Hall, pp. 69-93.

1994 Hartwig, WC Book Review: The Cambridge Encyclopaedia of Human Evolution. Evolutionary Anthropology 2:147-149.

1994 Hartwig, WC Embryology in the context of clinical gross anatomy: a review of current texts. Clinical Anatomy 7:108-109.

1994 Hartwig, WC Setting the Future Agenda for Research on Neotropical Primates: A Symposium in Honor of Warren Kinzey. Evolutionary Anthropology 2:189-192.

1994 Hartwig, WC Book Review: Origins of Anatomically Modern Humans (MH Nitecki and DV Nitecki, eds). Quarterly Review of Biology 69:562-563.

1993 Hartwig, WC Comparative Morphology, Ontogeny and Phylogeny of the Platyrrhine Cranium. PhD Dissertation, University of California, Berkeley.

1993 Hartwig, WC and LL Sadler Visualization and the future of physical anthropology. In Milestones in Human Evolution. Perspectives in Human Evolution, Volume 6 (A. Almquist and A. Manyak, eds.). Chicago: Waveland Press, pp. 187-222.

1992 Hartwig, WC Book Review: Jablonski's Dictionary of Syndromes and Eponymic Diseases. Clinical Anatomy 5:492-493.

1991 Hartwig, WC Fractal analysis of sagittal suture morphology. Journal of Morphology 210:289-298.

1991 Hartwig, WC Book Review: Pernkopf's Anatomy, 3rd ed. (W. Platzer, ed.) Clinical Anatomy 4:63-64.

1991 Rosenberger, AL, T Setoguchi and WC Hartwig Laventiana annectens: new fossil evidence for the origin of callitrichine New World monkeys. Proceedings of the National Academy of Sciences 88:2137-2140.

1991 Rosenberger, AL, WC Hartwig, M Takai, T Setoguchi and N Shigehara Dental variability in Saimiri and the taxonomic status of Neosaimiri fieldsi, an early squirrel monkey from Colombia, South America. International Journal of Primatology 12:291-301.

1991 Hartwig, WC, T Setoguchi and AL Rosenberger The emergence of modern platyrrhine genera in the Miocene of Colombia. Proceedings of the XIIIth Congress of the International Primatological Society. Amsterdam: Elsevier, pp. 525-526.

1991 Rosenberger, AL, WC Hartwig and RG Wolff Szalatavus attricuspis, an early platyrrhine primate. Folia Primatologica 56:225-233.

1990 Hartwig, WC Dental metrics and the radiation of New World monkeys: A preliminary analysis. Kroeber Anthropological Society Papers 71/72:57-66.

1990 Hartwig, WC Book Review: The Anatomy and Biology of the Human Skeleton, by D. Gentry Steele and Claud Bramblett. Clinical Anatomy 3:151-153.


Popular

1999 Hartwig, WC Under Brazil: Mapping the Caves of Ancient Amazonia. Mercator’s World 5:24-31

2005 Hartwig, WC The intelligent design of scientific inquiry. Napa Valley Register, Guest Commentary, 11 July 2005.

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Research Grants

1988 $800.00 Paleontological Survey of Middle Miocene Deposits at La Venta, Colombia. Graduate Division, University of California, Berkeley
1990 $500.00 Paleontological Survey of Tertiary Deposits along the Cordillera Merida and Orinoco Basin, Venezuela. Graduate Division, University of California, Berkeley
1991 $1100.00 Paleontological Survey of Tertiary Deposits along the Cordillera Merida and Orinoco Basin, Venezuela. Explorers Club
1991 $500.00 Paleontological Survey of Tertiary Deposits along the Cordillera Merida and Orinoco Basin, Venezuela. Department of Anthropology, University of California, Berkeley
1992 $375.00 Biogeography of New World Monkeys and the Pleistocene Caves of Lagoa Santa, Brazil. Sigma XI Society of Scientific Research
1992 $14,900.00 An Interactive Image Archive System for the Study of Prehistory. National Geographic Society
1993 $1500.00 Workshop on Imaging Research in Physical Anthropology. LSB Leakey Foundation
1994 $3450.00 New Large-bodied Primates in the Pleistocene of Brazil. LSB Leakey Foundation
1998 $18,920.00 Biogeography and Biodiversity of Pleistocene Mammals in Southeastern
Amazonia. National Geographic Society
2004 $11,910.00 Primate Origins and Paleocommunities (POPCOM): a Global Field Test of the
Early Origins Hypothesis. Touro University – California.

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